CONODONT BIOSTRATIGRAPHY AND LITHOSTRATIGRAPHY OF THE LATE GIVETIAN–EARLY FRASNIAN INTERVAL IN THE ARPI SECTION (CENTRAL ARMENIA)

Authors

DOI:

https://doi.org/10.54103/2039-4942/29903

Keywords:

Ancyrodella; conodonts; Givetian; Frasnian; Gondwana; Lesser Caucasus.

Abstract

The Givetian–Frasnian (G–F) transition marks a time of significant palaeoenvironmental perturbations and biotic turnover, yet its stratigraphic record in the southern Caucasus remains poorly constrained. This study presents the first integrated examination of conodont biostratigraphy and lithofacies across the G–F transitional interval in Armenia, based on material from the Arpi section. The section yielded a relatively diverse and abundant conodont assemblage, with taxonomic assessment allowing to identify eleven species and subspecies assigned to genera Polygnathus, Icriodus, and Ancyrodella. The first occurrences of Ancyrodella rotundiloba s.s., A. pristina, and A. binodosa in the upper part of the section enable correlation with the upper FZ1–FZ2 zones of the Frasnian conodont zonation, constraining this interval to the lowermost Frasnian. The absence of Ancyrodella in the underlying strata, coupled with the presence of species such as Polygnathus dubius, Icriodus cedarensis, and I. excavatus, suggests that the lower to middle part of the section corresponds to the upper Givetian. This interval is further correlated with the Upper hermanni to norrisi zones. Lithostratigraphic observations and microfacies analyses indicate that the sedimentary sequence of the section accumulated in a ramp setting. Moreover, a shallowing trend is observed, from relatively deeper, low-energy black shale facies in the late Givetian to more oxygenated, proximal carbonate facies in the early Frasnian.

Downloads

Download data is not yet available.

References

tian to basal Frasnian conodont fauna from the Tafilalt (Anti-Atlas, southern Morocco). Geologica Quarterly, 51: 345-374.

Abrahamyan M.S. (1959) - Novyye vidy brakhiopod iz verkhnego devona Armenii [New brachiopod species from the Upper Devonian of Armenia]. Izvestiya Akademii Nauk Armyanskoy SSR, 12: 4-10 [in Russian and in Armenian].

Abrahamyan M.S. (1964) - Karbon [Carboniferous]. In: Mkrtchian S.S., Vardaniants L.A., Gabrielian A.A, Maghakian I.G. & Paffenholz C.N. (Eds) - Geologiya Armyanskoy SSR [Geology of the Armenian SSR], 2: 96-118. Yerevan, Akademiya Nauk Armyanskoy SSR. [in Russian]

Abrahamyan M.S., Arakelyan R.A. & Azizbekov Sh. A. (1973) - Malyy Kavkaz (Yuzhnoe Zakavkaz’ya) [Lesser Caucasus (South Transcaucasia)]. In: Rzhonsnitskaya M.A. (Ed.) - Kavkazskaya Geosinklinal’naya Oblast i Predkavkaz’e [Caucasian geosynclinal region and Ciscaucasia]. In: Nalivkin D.V., Rzhonsnitskaya M.A. & Markovski B.P. (Eds) 1: 210-219. Stratigrafiya SSSR, Devonskaya Sistema Nedra, Moscow [in Russian].

Arakelyan R.A. (1952) - Stratigrafiya paleozoyskikh otlozheniy yugo-zapadnoy Armenii i prilegayushchikh chastey Nakhichevanskoy ASSR [Stratigraphy of the Palaeozoic deposits of southwestern Armenia and the adjacent parts of the Nakhichevan ASSR]. Yerevan, Akademiya Nauk Armyanskoy SSR, Institut Geologicheskikh Nauk, Yerevan, 142 pp. [in Russian].

Arakelyan R.A. (1964) - Devon [The Devonian]. In: Mkrtchian S.S., Vardaniants L.A., Gabrielian A.A., Maghakian I.G. & Paffenholz C.N. (Eds) - Geologiya Armyanskoy SSR [Geology of the Armenian SSR], 2: 46-96. Akademiya Nauk Armyanskoy SSR, Yerevan [in Russian].

Aristov V.A. (1994) - Konodonty devona-nizhnego karbona Evrazii: soobshhestva, zonalnoye raschleneniye, korrelyatsiya raznofatsialnykh otlozheniy [Conodonts from the Devonian and Lower Carboniferous of Eurasia: communities, zonation and correlation of heterofacial deposits]. Trudy Geologicheskogo Instituta Rossiyskoy Akademii Nauk, 484: 1-193 [in Russian].

Ashouri A.R. (2004) - Late Devonian and Middle–Late Devonian conodonts from eastern and northern Iran. Revista Española de Micropaleontología, 36: 355-365.

Ashouri A.R. (2006) - Middle Devonian–Early Carboniferous conodont faunas from the Khoshyeilagh Formation, Alborz Mountains, North Iran. Journal of Sciences, Islamic Republic of Iran, 17: 53-65.

Becker R.T., Gradstein F.M. & Hammer O. (2012) - The Devonian Period. In: Gradstein F.M., Ogg J.G., Schmitz M.D. & Ogg G.M. (Eds) - The Geologic Time Scale: 559-601, Elsevier, Amsterdam. DOI: https://doi.org/10.1016/B978-0-444-59425-9.00022-6

Becker R.T., Aboussalam Z.S., El Hassani A., Eichholt S. & Helling S. (2020) - The Devonian of the Oued Cherrat Zone (Western Meseta) – review and new data. Frontiers in Science and Engineering, Earth, Water and Oceans, Environmental Sciences, 10: 27-85.

Bensaid M., Bultynck P., Sartenaer P., Walliser O.H. & Ziegler W. (1985) - The Givetian–Frasnian boundary in pre-Sahara Morocco. Courier Forschungsinstitut Senckenberg, 75: 287-300.

Bertola C., Boulvain F., Da Silva A.-C. & Poty E. (2013) - Sedimentology and magnetic susceptibility of Mississippian (Tournaisian) carbonate sections in Belgium. Bulletin of Geosciences, 88: 69-82. DOI: https://doi.org/10.3140/bull.geosci.1339

Braun W.K., Norris A.W. & Uyeno T.T. (1988) - Late Givetian to Early Frasnian biostratigraphy of western Canada: The Slave Point–Waterways boundary and related events. In: McMillan, N.J., Embry A.F. & Glass D. J. (Eds) - Devonian of the World. 3: Paleontology, Paleoecology, Biostratigraphy. Proceedings of the Second International Symposium on the Devonian System, Memoirs 14: 93-111. Canadian Society of Petroleum Geologists, Calgary.

Buggisch W. & Joachimski M.M. (2006) - Carbon isotope stratigraphy of the Devonian of Central and Southern Europe. Palaeogeography, Palaeoclimatology, Palaeoecology, 240: 68-88. DOI: https://doi.org/10.1016/j.palaeo.2006.03.046

Bultynck P. (1986) - Accuracy and reliability of conodont zones: the Polygnathus asymmetricus “Zone” and the Givetian–Frasnian boundary. Bulletin de l’Institut royal des Sciences naturelles de Belgique, Sciences de la Terre, 56: 269-280.

Burchette T.P. & Wright V.P. (1992) - Carbonate ramp depositional systems. Sedimentary Geology, 79: 3-57. DOI: https://doi.org/10.1016/0037-0738(92)90003-A

Casier J.G. & Préat A. (2009) - Late Givetian to Middle Frasnian ostracods from Nismes (Dinant Synclinorium, Belgium) and their lithological context. Bulletin de l’Institut Royal des Sciences Naturelles de Belgique, Sciences de la Terre, 79: 87-115.

Casier J.G., Devleeschouwer X., Maillet S., Petitclerc E. & Préat A. (2013) - Ostracods and rock facies across the Givetian/Frasnian boundary interval in the Sourd d’Ave section at Ave-et-Auffe (Dinant Synclinorium, Ardenne, Belgium). Bulletin of Geosciences, 88: 241-264. DOI: https://doi.org/10.3140/bull.geosci.1340

Copper P. (1967) - Adaptations and life habits of Devonian atrypid brachiopods. Palaeogeography, Palaeoclimatology, Palaeoecology, 3: 363-379. DOI: https://doi.org/10.1016/0031-0182(67)90025-9

Corradini C., Henderson C., Barrick J.E. & Ferretti A. (2026) - Conodonts in Biostratigraphy. A 300-million-years long journey through geologic time. Newsletters on Stratigraphy, 59: 77-116. DOI: https://doi.org/10.1127/nos/2024/0822

Devleeschouwer X., Petitclerc E., Spassov S. & Préat A. (2010) - The Givetian–Frasnian boundary at Nismes parastratotype (Belgium): The magnetic susceptibility signal controlled by ferromagnetic minerals. Geologica Belgica, 13: 351-366.

Ebert J. (1993) - Globale Events im Grenz-Bereich Mittel-/Ober-Devon. Gottinger Arbeiten zur Geologie und Paläontologie 59: 1-106. University of Göttingen, Göttingen.

Feist R. & Klapper G. (1985) - Stratigraphy and conodonts in pelagic sequences across the Middle–Upper Devonian boundary, Montagne Noire, France. Palaeontographica Abteilung A, 188: 1-18.

Flügel E. (2004) - Microfacies of carbonate rocks. Springer-Verlag, Berlin, 976 pp. DOI: https://doi.org/10.1007/978-3-662-08726-8

Gholamalian H., Hairapetian V., Barfehei N., Mangelian S. & Faridi P. (2013) - Givetian–Frasnian boundary conodonts from Kerman Province, Central Iran. Rivista Italiana di Paleontologia e Stratigrafia, 119: 141-158.

Gholamalian H., Hairapetian V. & Poosti M. (2022) - Middle–Late Devonian boundary conodonts from Anarak and Sar-Ashk sections, Central Iran. Turkish Journal of Earth Sciences, 31: 71-84.

Gouwy S., Hayduckiewicz J. & Bultynck P. (2007) - Conodont-based graphic correlation of upper Givetian–Frasnian sections of the Eastern Anti-Atlas (Morocco). Geologica Quarterly, 51: 375-392.

Grechishnikova I.A., Levitskii E.S., Polozhikhina A.I. & Feliks V.P. (1983). - Kratkaya kharakteristika opornogo razreza srednego devona u sela Danzik (Nakhichevanskaya ASSR). [Brief characteristics of the reference section of the Middle Devonian near the village of Danzik (Nakhchivan ASSR)]. Izvestiya Vysshikh Uchebnykh Zavedeniy, Geologiya i Razvedka, 11: 98-105.

Herbig H.-G. & Weber H.M. (1996) - Facies and stromatoporoid biostromes in the Strunian (Latest Devonian) of the Aachen region, Germany. In: Reitner J., Neuweiler F. & Gunkel F. (Eds) - Global and regional controls on biogenic sedimentation. I. Reef evolution 2: 359-364. Göttinger Arbeiten zur Geologie und Paläontologie, Sonderband, Göttingen.

Hinde G.J. (1879) - On conodonts from the Chazy and Cincinnati Group of the Cambro–Silurian, and from the Hamilton and Genesse-Shale divisions of the Devonian, in Canada and the United States. Quarterly Journal of the Geological Society of London, 35: 351-369. DOI: https://doi.org/10.1144/GSL.JGS.1879.035.01-04.23

House M.R. (1985) - Correlation of mid-Palaeozoic ammonoid evolutionary events with global sedimentary perturbations. Nature, 313: 17-22. DOI: https://doi.org/10.1038/313017a0

House M.R. (2002) - Strength, timing, setting and cause of mid-Palaeozoic extinctions. Palaeogeography, Palaeoclimatology, Palaeoecology, 181: 5-25. DOI: https://doi.org/10.1016/S0031-0182(01)00471-0

Hua L., Ma X. & Wei L. (2009) - A Middle–Upper Devonian boundary section in the open-platform, platform-margin facies of Guilin, South China. Acta Geologica Sinica, 83: 524-534. DOI: https://doi.org/10.1111/j.1755-6724.2009.00048.x

Khalymbadzha V.G. & Chernysheva N.G. (1970) - Konodonty rod Ancyrodella iz Devonskikh otlozheniy Volgo-Kamskogo kraya i ikh stratigraficheskoe znachenie [Conodonts of the genus Ancyrodella from the Devonian deposits of the Volga-Kama region and their stratigraphic significance]. In: Biostratigrafiya I Paleontologiya Paleozoyskikh otlozheniy vostoka Russkoy Platformy I zapadnogo priural’ya [Biostratigraphy and paleontology of the Palaeozoic deposits of the Eastern Russian Platform and the Western Urals]. 1: 81-103. Izdatel’stvo Kazanskogo Universiteta Vypusk [in Russian].

Klapper G. (1985) - Sequence in the conodont genus Ancyrodella in the Lower asymmetricus Zone (earliest Frasnian, Upper Devonian) of the Montagne Noire, France. Palaeontographica, Abteilung A, 188: 19-34.

Klapper G. (1989) - The Montaigne Noire Frasnian (Upper Devonian) Conodont Succession. In: McMillan N.J., Embry D.J. & Glass D.J. (Eds) - Devonian of the World, Proceedings of the Second International Symposium on the Devonian System, 14: 449-468. Canadian Society of Petroleum Geologists.

Klapper G. (2000) - Species of Spathognathodontidae and Polygnathidae (Conodonta) in the recognition of Upper Devonian stage boundaries. Courier Forschungsinstitut Senckenberg, 220: 153-159.

Klapper G. (2021) - Revision of the Frasnian Late Devonian conodont genus Ancyrodella. Bulletin of Geosciences, 96: 295-325. DOI: https://doi.org/10.3140/bull.geosci.1828

Klapper G. & Johnson J.G. (1990) - Revisions of Middle Devonian conodont zones. In: Johnson J.G. (Ed.) - Lower and Middle Devonian brachiopod-dominated communities of Nevada, and their position in a biofacies-province-realm model. Journal of Paleontology, 64: 934–941. DOI: https://doi.org/10.1017/S002233600002000X

Klapper G. & Kirchgasser W.T. (2016) - Frasnian Late Devonian conodont biostratigraphy in New York: Graphic correlation and taxonomy. Journal of Paleontology, 90: 525-554. DOI: https://doi.org/10.1017/jpa.2015.70

Klapper G. & Ziegler W. (1979) - Devonian conodont biostratigraphy. Special Papers in Palaeontology, 23: 199-224.

Klapper G., Feist R. & House M.R. (1987) - Decision on the boundary stratotype for the Middle/Upper Devonian Series boundary. Episodes, 10: 97-101. DOI: https://doi.org/10.18814/epiiugs/1987/v10i2/004

Kononova L.I., Alekseev A.S., Barskov I.S & REIMERS A.N. (1996) - New species of Polygnathid assemblages from the Frasnian Stage of the Moscow Syneclise. Paleontologicheskii Zhurnal 3: 94-99.

Liao J.C. & Valenzuela-Ríos J.I. (2008) - Givetian and early Frasnian conodonts from the Compte section (Middle–Upper Devonian, Spanish Central Pyrenees). Geologica Quarterly, 52: 1-18.

Liao J.-C. & Valenzuela-Ríos J.I. (2012) - Upper Givetian and Frasnian (Middle and Upper Devonian) conodonts from Ampriú (Aragonian Pyrenees, Spain): Global correlations and palaeogeographic relations. Palaeontology, 55: 819-842. DOI: https://doi.org/10.1111/j.1475-4983.2012.01150.x

Liao J.-C. & Valenzuela-Ríos J.I. (2013) - The Middle and Upper Devonian conodont sequence from La Guardia d’Ares sections (Spanish Central Pyrenees). Bulletin of Geosciences, 88: 339-368. DOI: https://doi.org/10.3140/bull.geosci.1348

Liao J.-C. & Valenzuela-Ríos J.I. (2017) - Givetian (Middle Devonian) historical bio- and chronostratigraphical subdivision based on conodonts. Lethaia, 50: 440-446. DOI: https://doi.org/10.1111/let.12192

Liao J.C., Girard C., ValenzuelaRíos J.I. & Feist R. (2019) - New conodont data from the Middle–Upper Devonian boundary stratotype at Col du Puech de la Suque (Montagne Noire, France). STRATI 2019, Third International Congress on Stratigraphy, Milano, p. 188.

Mabille C. & Boulvain F. (2007) - Sedimentology and magnetic susceptibility of the Upper Eifelian –Lower Givetian (Middle Devonian) in southwestern Belgium: Insights into carbonate platform initiation. In Álvaro J.J., Aretz M., Boulvain F., Munnecke A., Vachard D. & Vennin E. (Eds) - Palaeozoic reefs and bioaccumulations. Climatic and evolutionary controls, 275: 109-124. Geological Society, Special Publications, London. DOI: https://doi.org/10.1144/GSL.SP.2007.275.01.08

Milanovsky E.E. (1968) - Noveyshaya tektonika Kavkaza [Neotectonics of the Caucasus]. Nedra, Moscow, 483 pp. [in Russian].

McGhee G.R., Clapham M.E., Sheehan P.M., Bottjer D.J. & Droser M.L. (2013) - A new ecological-severity ranking of major Phanerozoic biodiversity crises. Palaeogeography, Palaeoclimatology, Palaeoecology, 370: 260-270. DOI: https://doi.org/10.1016/j.palaeo.2012.12.019

Narkiewicz K. & Bultynck P. (2007) - Conodont biostratigraphy of shallow marine Givetian deposits from the Radom-Lublin area, SE Poland. Geological Quarterly, 51: 419-442.

Narkiewicz K. & Bultynck P. (2010) - The Upper Givetian (Middle Devonian) subterminus conodont zone in North America, Europe, and North Africa. Journal of Paleontology, 84: 588-625. DOI: https://doi.org/10.1666/10-029.1

Nikogosian I.K., Bracco Gartner A.J., Mason P.R., van Hinsbergen D.J.J., Kuiper K.F., Kirscher U., Matveev S., Grigoryan A., Grigoryan E., Israyelyan A., van Bergen M.J., Koornneef J.M., Wijbrans J.R., Davies G.R. & Meliksetian K. (2023) - The South Armenian Block: Gondwanan origin and Tethyan evolution in space and time. Gondwana Research, 121: 168-195. DOI: https://doi.org/10.1016/j.gr.2023.03.023

Ovnatanova N.S. & Aristov V.A. (1984) - Biostratigrafiya verkhnedevonskikh otlozheniy Zakavkaz’ya po konodontam [Biostratigraphy of the Upper Devonian deposits of Transcaucasia based on conodonts]. Izvestiya Akademii Nauk SSSR, Seriya Geologicheskaya, 5: 61-68 [in Russian].

Özkan R., Nazik A., Munnecke A., Demiray D.G., Schindler E., Özbek T.A., Zor E.Ş., Yılmaz İ., Brocke R., Sancay R.H., Wilde V. & Yalçın M.N. (2019) - Givetian–Frasnian (Middle/Upper Devonian) transition in the eastern Taurides, Turkey. Turkish Journal of Earth Sciences, 28: 207-231. DOI: https://doi.org/10.3906/yer-1804-20

Qie W., Liang K. & Königshof P. (2019) - Devonian palaeoecosystems and palaeoenvironments of South China. Palaeobiodiversity and Palaeoenvironments, 99: 1-5. DOI: https://doi.org/10.1007/s12549-018-00370-3

Racki G. (1993) - Evolution of the bank to reef complex in the Devonian of the Holy Cross Mountains. Acta Palaeontologica Polonica, 37: 87-182.

Racki G. & Wrzolek T. (1989) - Middle–Upper Devonian boundary: Ambiguous reality of its stratotype. Courier Forschungsinstitut Senckenberg, 110: 231-236.

Read J.E. (1985) - Carbonate platform facies models. Bulletin of the American Association of Petroleum Geologists, 69: 1-21. DOI: https://doi.org/10.1306/AD461B79-16F7-11D7-8645000102C1865D

Sandberg C.A., Ziegler W. & Bultynck P. (1989) - New standard conodont zones and early Ancyrodella phylogeny across the Middle–Upper Devonian boundary. Courier Forschungsinstitut Senckenberg, 110: 195-230.

Sepkoski J.J. Jr., Bambach R.K. & Droser M.L. (1991) - Secular changes in Phanerozoic event bedding and biological overprint. In: Einsele G., Ricken W. & Seilacher A. (Eds) - Cycles and events in stratigraphy: 298-312. Berlin, Springer.

Serobyan V., Grigoryan A., Mottequin B., Mayilyan R., Crônier C. & Danelian T. (2019) - Biostratigraphy of the Upper Devonian trigonirhynchiid brachiopods (Rhynchonellida) from Armenia. Proceedings of the National Academy of Sciences of the Republic of Armenia, Earth Sciences, 72: 3-18.

Serobyan V., Danelian T., Hairapetian V., Grigoryan A., Crônier C., Randon C. & Mottequin B. (2023) - Frasnian (Upper Devonian) brachiopods from Armenia: Biostratigraphic and paleobiogeographic implications. Rivista Italiana di Paleontologia e Stratigrafia, 129: 373-409. DOI: https://doi.org/10.54103/2039-4942/19826

Sosson M., Rolland Y., Müller C., Danelian T., Melkonyan R., Kekelia S., Adamia S., Babazadeh V., Kangarli T., Avagyan A., Galoyan G. & Mosar J. (2010) - Subductions, obduction and collision in the Lesser Caucasus (Armenia, Azerbaijan, Georgia), new insights. In: Sosson M., Kaymakci N., Stephenson E.A., Bergerat F. & Starostenko V. (Eds) - Sedimentary Basin Tectonics from the Black Sea and Caucasus to the Arabian Platform 340: 329-352. Geological Society, London. DOI: https://doi.org/10.1144/SP340.14

Stauffer C.R. (1938) - Conodonts of the Olentangy Shale. Journal of Paleontology, 12: 411-443.

Stichling S., Becker R.T., Hartenfels S., Aboussalam Z.S. & May A. (2022) - Drowning, extinction, and subsequent facies development of the Devonian Hönne Valley Reef (northern Rhenish Massif, Germany). Palaeobiodiversity and Palaeoenvironments, 102: 629-696. DOI: https://doi.org/10.1007/s12549-022-00539-x

Uyeno T.T. (1967) - Conodont zonation, Waterways Formation (Upper Devonian), northeastern and central Alberta. Geological Survey of Canada Paper, 67: 1-21. DOI: https://doi.org/10.4095/100913

Walliser O.H. (1996) - Global events in the Devonian and Carboniferous. In: Walliser O.H. (Ed.) - Global Events and Event Stratigraphy: 225-250. Springer-Verlag, Berlin. DOI: https://doi.org/10.1007/978-3-642-79634-0_11

Weddige K. (1984) - Zur Stratigraphie und paläogeographie des Devons und Karbons von NE Iran. Senckenbergiana lethaea, 65: 179-223.

Wehrmann A., Yılmaz I., Yalçın M.N., Wilde V., Schindler E., Weddige K., Saydam Demiray G., Özkan R., Nazik A. & Nalcıoğlu G. (2010) - Devonian shallow-water sequences from the north Gondwana coastal margin (central and eastern Taurides, Turkey). Gondwana Research, 17: 546-560. DOI: https://doi.org/10.1016/j.gr.2009.09.011

Ziegler W. & Klapper G. (1982) - The disparilis conodont Zone, the proposed level for the Middle–Upper Devonian boundary. Courier Forschungsinstitut Senckenberg, 55: 463-492.

Ziegler W. & Sandberg C.A. (1996) - Reflexions on Frasnian and Famennian Stage boundary decisions as a guide to future deliberations. Newsletters on Stratigraphy, 33: 157-180. DOI: https://doi.org/10.1127/nos/33/1995/157

Ziegler W. & Sandberg C.A. (2000) - Utility of palmatolepids and icriodontids in recognizing Upper Devonian series, stage, and possible substage boundaries. Courier Forschungsinstitut Senckenberg, 225: 335-347.

Ziegler W., Klapper G. & Johnson J.G. (1976) - Redefinition and subdivision of the varcus-Zone (Conodonts, Middle–?Upper Devonian) in Europe and North America. Geologica et Palaeontologica, 10: 109-140.

Downloads

Published

09-09-2026

How to Cite

SEROBYAN, V. (2026) “CONODONT BIOSTRATIGRAPHY AND LITHOSTRATIGRAPHY OF THE LATE GIVETIAN–EARLY FRASNIAN INTERVAL IN THE ARPI SECTION (CENTRAL ARMENIA)”, RIVISTA ITALIANA DI PALEONTOLOGIA E STRATIGRAFIA , 132(3). doi: 10.54103/2039-4942/29903.

Issue

Section

Articles
Received 2025-10-12
Accepted 2026-08-05
Published 2026-09-09

Funding data